Wow, that was a fascinating read and makes complete sense. Developing language moved to abstract concepts because words had to represent things and brains that supported that had an advantage
The ethical concern aspect making it difficult to study is unfortunate - would be curious if elephants or orca could be shown to have evidence of concept neurons. Elephants have been shown to recognize accents/language in humans and avoided humans who are associated with elephant hunting tribes. Orcas also are frequently observed teaching each other skills or behaviors
Also would be impossible to tell, but I wonder if these existed in other human species, like Neanderthal